Pseudocaeciliinae
Belongs within: Pseudocaecilioidea.
The Pseudocaeciliinae are a group of barklice with two-segmented tarsi, known predominantly from south-east Asia and the Pacific (New & Lienhard 2007).
<==Pseudocaeciliinae [Meplerinae, Pseudoscottiinae] NL07
|--*Allopsocus marginalis Banks 1920 NL07
|--Scottiella NL07
|--Platyocaecilius Li 2002 L02b
| `--*P. parallelivenius Li 2002 L02b
|--Fashengocaecilius Lienhard 2003 L03 [=Calocaecilius Li 2002 non Mockford 1974 L02b]
| `--*F. biaduncus (Li 1992) [=Allocaecilius biaduncus, *Calocaecilius biaduncus] L03
|--Phyllocaecilius Li 2002 L02b
| `--*P. atrichus Li 2002 L02b
|--Chorocaecilius Li 2000 L02a
| |--C. brunellus (Tillyard 1923) (see below for synonymy) SN08
| `--C. decemdentatus Li 2000 L02a
|--Phallocaecilius Lee & Thornton 1967 L02b
| |--*P. hirsutus (Thornton 1961) [=Pseudocaecilius hirsutus] NL07
| `--P. sentosus Li 1999 L02a
|--Thelocaecilius Li 2002 L02b
| |--*T. mecokeratus Li 2002 L02b
| `--T. papillatus (Li 1995) [=Heterocaecilius papillatus] L02b
|--Neocaecilius Li 2000 L02a
| |--*N. campanula [=Heterocaecilius campanula] NL07
| |--N. granulatus (Lee & New 1992) NL07
| |--N. mutatus (Lee & New 1992) NL07
| `--N. qianshanensis Li 2000 L02a
|--Mesocaecilius Okamoto 1910 NL07, L02a [Mesocaeciliinae]
| |--*M. quadrimaculatus Okamoto 1910 L02a
| |--M. bellus Li 2002 L02b
| |--M. decorosus Li 2002 L02b
| |--M. elegans Li 1997 L02b
| |--M. euryopterus Li 2002 L02b
| |--M. lepidus Li 2002 L02b
| `--M. paraornatus (New 1977) [=Pseudocaecilius paraornatus] L02b
|--Ophiodopelma Enderlein 1908 L02a
| |--*O. ornatipenne Enderlein 1908 NL07, L02a
| |--O. anocella Li 2002 L02b
| |--O. fluctosa Li 1995 L02a
| |--O. glyptocephalum Lienhard 1985 NL07
| |--O. hieroglyphicum (Enderlein 1903) NL07
| |--O. multipunctatum (Hagen 1859) NL07
| |--O. permaculatum Lee & Thornton 1967 NL07
| |--O. pictipenna Lee & Thornton 1967 NL07
| |--O. polyspila Li 1993 L02b
| `--O. semiceps Lee & Thornton 1967 NL07
|--Levucaecilius Smithers 1995 NL07, S95
| |--*L. greenwoodi [=Pseudocaecilius greenwoodi, Orbocaecilius greenwoodi] NL07
| |--L. albicrus (Thornton 1981) [=Heterocaecilius albicrus] S95
| |--L. apicalis (Thornton 1981) [=Heterocaecilius apicalis] S95
| |--L. dardanus (Lee & Thornton 1967) S95 [=Heterocaecilius dardanus NL07, Neocaecilius dardanus NL07]
| |--L. hilli Smithers 1995 NL07
| |--L. panicus (Lee & Thornton 1967) [=Heterocaecilius panicus] S95
| |--L. pictus (Thornton 1981) [=Heterocaecilius pictus] S95
| |--L. simplex (Lee & Thornton 1967) NL07, L02a [=Heterocaecilius simplex NL07]
| |--L. veitchi [=Heterocaecilius veitchi] S95
| `--L. volatus (Lee & Thornton 1967) [=Heterocaecilius volatus] S95
|--Lobocaecilius Lee & Thornton 1967 L02a
| |--*L. cynara Lee & Thornton 1967 L02a
| |--L. arcuatus Vaughan, Thornton & New 1989 NL07
| |--L. bifurcus Li 1993 L02a
| |--L. drepanus New & Thornton 1976 NL07
| |--L. ligur Lee & New 1992 NL07
| |--L. longifurca Lee & Thornton 1967 NL07
| |--L. pahangensis Lee & New 1992 NL07
| |--L. perductivirgus Li 2002 L02b
| |--L. pilosus Lee & New 1992 NL07
| |--L. quadripartitus Li 1995 L02b
| |--L. thorntoni Lee & New 1992 NL07
| `--L. viridis Lee & New 1992 NL07
|--Allocaecilius Lee & Thornton 1967 L02a
| |--*A. heterothrax Lee & Thornton 1967 L02a
| |--A. albidorsualis Li 1993 L02a
| |--A. arcuatus Li 2002 L02b
| |--A. elongatus Lee & Thornton 1967 NL07
| |--A. fusciceps Lee & Thornton 1967 NL07
| |--A. gombakensis Lee & New 1992 NL07
| |--A. guangdongicus Li 1993 L02a
| |--A. maculatus Lee & Thornton 1967 NL07
| |--A. selangorensis New & Thornton 1975 NL07
| |--A. sinensis Lee & Thornton 1967 L02a
| |--A. smithersi Lee & New 1992 NL07
| `--A. tenuilongus Li 1995 L02a
|--Mepleres Enderlein 1926 SN08 [incl. Meniscopsocus Li 1993 L02b, Pseudoscottiella Badonnel 1946 L02b]
| |--*M. maeandricus Enderlein 1926 SN08
| |--M. cornutus (Lee & New 1992) NL07 [=Pseudoscottiella cornuta L02b]
| |--‘Pseudoscottiella’ decolor Badonnel 1955 B55
| |--‘Pseudoscottiella’ hyalina Badonnel 1955 B55
| |--‘Pseudoscottiella’ immaculata Badonnel 1955 B55
| |--M. indicus (Lee & New 1992) NL07
| |--M. limbatus (Enderlein 1908) L02a
| |--M. longicellus Li 2002 L02b
| |--M. longitudinalis Li 2002 [=M. longitudianlis] L03
| |--‘Pseudoscottiella’ maculata Broadhead & Richard 1982 L02b
| |--‘*Pseudoscottiella’ megops NL07
| |--M. nigroguttatus (Karny 1925) NL07 [=Pseudoscottiella nigraguttata L02b]
| |--M. ocellatus Li 2002 L02b
| |--M. oresbius Li 2002 L02b
| |--M. parvicellus Li 2002 L02b
| |--M. pisiformis (Lee & New 1992) NL07
| |--M. plenimacularis Li 2002 L02b [=Pseudoscottiella plenimacularis L02a]
| |--M. proboscideus Li 2002 L02b
| |--M. procurrens Li 2002 L02b
| |--M. radialis (Thornton 1984) NL07
| |--M. raphalis (Lee & New 1992) NL07
| |--M. rotundatus (New 1974) [=Pseudoscottiella rotundata, Meniscopsocus rotundatus] SN08
| |--M. similis (Lee & New 1992) NL07
| |--M. sinicus (Li 1993) L02b [=*Meniscopsocus sinicus NL07]
| |--M. solenocerus (Lee & New 1992) NL07
| |--M. suella (Thornton 1984) NL07 [=Pseudoscottiella suella L02b]
| |--M. tanei (Smithers 1977) [=Pseudoscottiella tanei, Meniscopsocus tanei] SN08
| |--M. transversus (Banks 1937) NL07
| |--‘Pseudoscottiella’ tuberculata Badonnel 1955 B55
| |--M. ulterior Li 2002 L02b
| |--M. unitus Li 2002 L02b
| |--M. urbanus (Vaughan, Thornton & New 1991) NL07
| |--M. vilus (Lee & New 1992) NL07
| |--‘Pseudoscottiella’ yeni New 1974 L02b
| `--M. yunnanicus Li 2002 L02b
|--Pseudocaecilius Enderlein 1903 Y02 [incl. Hageniella Enderlein 1903 (preoc.) L02a]
| |--P. citricola (Ashmead 1879) NL07 (see below for synonymy)
| |--P. angustus (Enderlein 1903) NL07
| |--P. bibulbus Li 1993 L03, L02a [=P. bibalbus (l. c.) L03]
| |--P. bicostatus Li 2002 L02b
| |--P. ceratocercus Li 2002 L02b
| |--P. claggi (Banks 1937) NL07
| |--P. clunialis Vaughan, Thornton & Neo 1991 NL07
| |--P. cornutus Lee & Thornton 1967 NL07
| |--P. danaus Lee & New 1992 NL07
| |--P. euryocercus Li 2002 [=P. eurocerus] L03
| |--P. formosanus (Banks 1937) L02a
| |--P. galactozonalis Li 2002 [=P. galactozonacis] L03
| |--P. helicoides Lee & Thornton 1967 NL07
| |--P. immaculatus Li 2002 L02b
| |--P. inaequalis (Banks 1916) NL07
| |--P. innotatus Banks 1937 NL07
| |--P. kagoshimensis Y02
| |--P. largicellus Li 2002 L02b
| |--P. machadoi Badonnel 1955 B55
| |--P. monotaeniatus Li 2002 [=P. monotaenitus, P. monotaenius] L03
| |--P. nitoris (Banks 1937) NL07
| |--P. octomaculatus (Li 1995) [=Heterocaecilius octomaculatus] L02b
| |--P. otiosus (Banks 1937) NL07
| |--P. papillaris Li 2002 L02b
| |--P. plagiozonalis Li 1995 L02a
| |--P. productus New & Thornton 1976 NL07
| |--P. pusillus (Banks 1931) NL07
| |--P. ranus Thornton 1984 NL07
| |--P. serratus Lee & New 1992 NL07
| |--P. setifer Vaughan, Thornton & New 1991 NL07
| |--P. sexdentatus Li 2002 L02b
| |--P. tahitiensis (Karny 1926) [=Epipsocus tahitiensis] M93
| |--P. tenellus Enderlein 1926 NL07
| |--P. testaceus Enderlein 1903 NL07
| |--P. undecimimaculatus Li 1995 L02b
| |--P. utricularis Li 1999 L02a
| |--P. venimaculatus Li 2002 L02b
| |--P. wellsae Lee & New 1992 NL07
| `--‘Psocus’ zonatus [=*Hageniella zonata] NL07
`--Heterocaecilius Lee & Thornton 1967 L02a [incl. Orbocaecilius Li 1999 L03, Obrocaecilius (l. c.) L03]
|--*H. minutus Lee & Thornton 1967 SN08 [=H. minotus NL07]
|--H. abacutidens Li 2002 L02b
|--H. analis Vaughan, Thornton & New 1991 NL07
|--H. anareolatus New 1993 NL07
|--H. anomalus (Thornton 1961) Y02, L02b [=Orbocaecilius anomolus L02b]
|--H. argutus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) argutus] L03
|--H. bicruris (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bicruris] L03
|--H. bidigitatus Li 1992 L02a [=Orbocaecilius bidigitatus L03]
|--H. bifarius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bifarius] L03
|--H. brachystigmus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) brachystigmus] L03
|--H. brevifurca Thornton 1984 NL07
|--H. challengerius Lee & New 1992 NL07
|--H. circulicellus Li 1995 L02b [=Orbocaecilius circulicellus L03]
|--H. dasoceratus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) dasoceratus] L03
|--H. deloneurous Li 2002 L02b
|--H. dentalis Li 2002 L02b
|--H. duodecidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) duodecidentus] L03
|--H. fasciatus Lee & Thornton 1967 NL07
|--H. flaviformis Lee & New 1992 NL07
|--H. fumosus Thornton 1984 NL07
|--H. fuscipalpus Lee & Thornton 1967 NL07
|--H. fuscus Y02
|--H. gressitti Lee & Thornton 1967 NL07
|--H. hepaticivenus Li 2002 L02b
|--H. hoplites Lee & New 1992 NL07
|--H. huananensis Li 2002 L02b
|--H. huaxiensis Li 2002 L02b
|--H. imparilis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) imparilis] L03
|--H. jinghongicus Li 1995 L02a
|--H. kobus Lee & Thornton 1967 NL07
|--H. lachlani (Enderlein 1903) [=Pseudocaecilius lachlani] SN08
|--H. leei New 1993 NL07
|--H. liangi Li 1991 L02b
|--H. loiaoi New 1991 L02a
|--H. longicornis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) longicornis] L03
|--H. longipenna Lee & Thornton 1967 NL07
|--H. longispinus Lee & Thorton 1967 NL07
|--H. maculans Li 1991 L02b
|--H. maculifornis (Thornton 1961) L02b [=Pseudocaecilius maculifrons L02a]
|--H. maculosus Lee & New 1992 NL07
|--H. marginatus Lee & New 1992 NL07
|--H. mecotrichus Li 2002 L02b
|--H. odontothelus Li 2002 L02b
|--H. orientalis Lee & New 1992 NL07
|--H. ornatus (Enderlein 1903) [=Pseudocaecilius ornatus] NL07
|--H. papillosus Vaughan, Thornton & New 1991 NL07
|--H. paulicellus (Li 1999) NL07, L02a [=*Orbocaecilius paulicellus L02a]
|--H. platokeratis Li 1993 L02a
|--H. pseudoanomalus Li 1993 L02b [=Obrocaecilius (l. c. for Orbocaecilius) pseudoanomalus L03]
|--H. quadrangulus Li 1992 L02b
|--H. reticulatus Lee & New 1992 NL07
|--H. rhadnus Thornton 1984 NL07
|--H. sedecimidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) sedecimidentus] L03
|--H. stenokeratis Li 1993 L02a
|--H. stiliger Lee & Thornton 1967 NL07
|--H. taipingensis Garcia Aldrete 1999 NL07
|--H. tartareus Li 2002 L02b
|--H. vulturius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) vulturius] L03
`--H. xihuicus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) xihuicus] L03
Chorocaecilius brunellus (Tillyard 1923) [=Caecilius brunellus, Heterocaecilius brunellus, Pseudocaecilius brunellus; incl. H. diogenes Lee & Thornton 1967, *Chorocaecilius diogenes] SN08
Pseudocaecilius citricola (Ashmead 1879) NL07 [=Psocus citricola M93; incl. Pse. elutus var. africanus Badonnel 1931 L02a, Elipsocus criniger Perkins 1899 M93, Kilauella criniger M93, Pseudocaecilius criniger M93, *Pse. elutus Enderlein 1903 NL07, L02b, Trichopsocus (*Pseudocaecilius) elutus L02a, Trichopsocus indicatus Navás 1934 M93, Pse. medianicostatus Li 1993 L02a, Caecilius pretiosus Banks 1920 M93, Pse. pretiosus M93, Pse. wolcotti Banks 1924 M93]
*Type species of generic name indicated
REFERENCES
[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.
[L02a] Li F. 2002a. Psocoptera of China vol. 1. Science Press: Beijing.
[L02b] Li F. 2002b. Psocoptera of China vol. 2. Science Press: Beijing.
[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.
[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.
[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.
[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.
[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.
[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.